External tufted cells have intrinsic burst firing and they are contacted directly by olfactory nerve (ON) terminals. They also fire in synchrony and contact inhibitory interneurons, namely periglomerular (PG) and short axon cells (SA), which in turn exhibit bursts of EPSPs. The PG cell dendrites ramify in one single glomerulus and thus might serve for intraglomerular inhibition via dendrodendritic interactions with ET tufted dendrites. In contrast, SA cell have neurites extending throughout several glomeruli and thus might serve for interglomerular inhibition.